Mechanism of Mineral Absorption in Plants: Active vs Passive Methods
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Plants absorb minerals as ions
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Plants absorb minerals from the soil in the
form of inorganic ions. Earlier it was thought
that the absorption of minerals takes place in
plants along with the absorption of water. In
fact, the process of mineral absorption and
water absorption are two separate processes.
In order to absorb any minerals from the soil,
it should be dissolved in the water.
•
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Minerals can be absorbed by both Passive
and Active Methods
•
There are two types of mineral absorptions
based on the involvement of metabolic
energy. They are (1) Passive minerals
absorption (2) Active minerals absorption.
•
(1). Passive Mineral Absorption
•
Passive mineral absorption is a passive process and
it does not require the expenditure of metabolic
energy. This type of mineral absorption occurs
along the concentration gradient by simple
diffusion.
•
(2). Active Mineral Absorption
•
Active mineral absorption is an active process and
thus it requires the expenditure of metabolic
energy (ATP). Active mineral absorption can occur
both along and against the concentration gradient
by osmosis or through special carrier proteins in
the plasma membrane.
Difference between Passive and Active Mineral Absorption
•
The exact mechanism of mineral absorption varies:
•
The exact mechanisms by which the plants absorb mineral
from the soil vary greatly for various types of minerals.
There are three main mechanisms of mineral absorption
processes in plants.
•
(1). Ion Exchange
•
(2). Carrier Concept
•
(3). Donnan’s Equilibrium
•
(1). Ion Exchange
•
In ion exchange process of mineral absorption, the ions
adsorbed on the surface of the root can exchange with the
ions of the same charge from the soil solution. For
example, H
+
ions adsorbed on the surface of root cells can
be exchanged with K
+
ions in the soil. Two theories have
been proposed to explain the mechanism of ion exchange
– (a) Contact exchange theory and (b) Carbonic acid
exchange theory.
•
(a). Contact Exchange Theory:
According to contact
exchange theory, the ions adsorbed on the surface of root
cells and clay particles are in continuous oscillation. When
the root and clay particles are in close contact with each
other, the oscillation radius of ions on the clay particles
may overlap with the ions adsorbed on the root cells.
When these oscillation radii overlap, there is a possibility of
spontaneous exchange of ions between clay particles and
root surface.
•
(b). Carbonic Acid Exchange Theory:
According to carbonic
acid exchange theory, the CO
2
produced in the root cells
due to respiration combine with water to form carbonic
acid (H
2
CO
3
). The carbonic acid is immediately dissociated
into H
+
and HCO
3
–
ions. The H
+
ions thus formed are
exchanged with cations on the clay particles.
•
(2). The Carrier Concept
•
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complex can move across the plasma membrane. When the carrier-
ion-complex reaches the inner surface of the membrane, they
release the ions into the lumen of the cell. After this, the carrier
protein will go back to the outer surface to accept new ions.
•
(3). Donnan Equilibrium
•
•
The Donnan equilibrium explains the accumulation of some mineral
ions inside the cell against the concentration gradient without the
expenditure of metabolic energy. According to this theory, inside
the root cells, there are some ions called Indiffusable Ions or Fixed
Ions, which do not diffuse outside through the plasma membrane.
The plasma membrane is permeable to other type of cations and
anions. If a cell has an accumulation of cations as fixed ions, such a
cell can absorb anions from the soil in order to maintain the
electrical potential balance. The absorption of anion in this case
causes an equilibrium of both cations and anion inside the cell. This
equilibrium is called Donnan equilibrium. By attaining the Donnan
equilibrium, the root cells can absorb any minerals in their ionic
form.
Plants absorb minerals from the soil in the form of inorganic ions through both passive and active methods. Active mineral absorption requires metabolic energy, whereas passive absorption occurs along the concentration gradient by simple diffusion. The roots play a vital role in mineral absorption due to their large surface area. Understanding the difference between passive and active absorption mechanisms is crucial for optimizing plant nutrition.
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Plant Physiology: by Dr. Manal Zbari Mechanism of Mineral Absorption in Plants lecture (2)
Mineral Absorption in Plants (The Mechanism of Active and Passive Absorption of Minerals in Plants) Plants absorb minerals as ions Plants absorb minerals from the soil in the form of inorganic ions. Earlier it was thought that the absorption of minerals takes place in plants along with the absorption of water. In fact, the process of mineral absorption and water absorption are two separate processes. In order to absorb any minerals from the soil, it should be dissolved in the water.
Plants absorb most of the minerals through the roots. The large surface area of roots and its ability to absorb minerals from the soil even in minor concentration makes the roots more efficient in mineral absorption than any other organs. In roots, the mineral absorption usually takes place through the meristematic region of the root tip.
Minerals can be absorbed by both Passive and Active Methods There are two types of mineral absorptions based on the involvement of metabolic energy. They are (1) Passive minerals absorption (2) Active minerals absorption.
(1). Passive Mineral Absorption Passive mineral absorption is a passive process and it does not require the expenditure of metabolic energy. This type of mineral absorption occurs along the concentration gradient by simple diffusion. (2). Active Mineral Absorption Active mineral absorption is an active process and thus it requires the expenditure of metabolic energy (ATP). Active mineral absorption can occur both along and against the concentration gradient by osmosis or through special carrier proteins in the plasma membrane.
Difference between Passive and Active Mineral Absorption Sl.No. Passive Absorption of Minerals Active Absorption of Minerals A passive, physical and An active, chemical and 1 spontaneous process. nonspontaneous process. Do not require energy. Energy required. 2 Movement of ions occurs Movement of ions occurs 3 along the concentration against the concentration gradient. gradient.
Movement of mineral ions Movement of mineral ions 4 proceeds towards the does not proceed towards equilibrium constant. the equilibrium constant. Does not cause salt Cause salt accumulation 5 accumulation in the cells. in the cells. Rate of mineral absorption Rate of mineral absorption 6 is independent of is dependent on respiration. respiration. Does not require specific Always require specific 7 carrier molecules. carrier proteins.
The efficiency of passive mineral The efficiency of 8 absorption is very less since the active mineral concentration of minerals inside the absorption is cell is usually 10,000 times more than comparatively that in the surrounding soil. efficient.
The exact mechanism of mineral absorption varies: The exact mechanisms by which the plants absorb mineral from the soil vary greatly for various types of minerals. There are three main mechanisms of mineral absorption processes in plants. (1). Ion Exchange (2). Carrier Concept (3). Donnan s Equilibrium (1). Ion Exchange In ion exchange process of mineral absorption, the ions adsorbed on the surface of the root can exchange with the ions of the same charge from the soil solution. For example, H+ions adsorbed on the surface of root cells can be exchanged with K+ions in the soil. Two theories have been proposed to explain the mechanism of ion exchange (a) Contact exchange theory and (b) Carbonic acid exchange theory.
(a). Contact Exchange Theory: According to contact exchange theory, the ions adsorbed on the surface of root cells and clay particles are in continuous oscillation. When the root and clay particles are in close contact with each other, the oscillation radius of ions on the clay particles may overlap with the ions adsorbed on the root cells. When these oscillation radii overlap, there is a possibility of spontaneous exchange of ions between clay particles and root surface. (b). Carbonic Acid Exchange Theory: According to carbonic acid exchange theory, the CO2produced in the root cells due to respiration combine with water to form carbonic acid (H2CO3). The carbonic acid is immediately dissociated into H+and HCO3 ions. The H+ions thus formed are exchanged with cations on the clay particles.
(2). The Carrier Concept According to this theory, the plasma membrane is completely impermeable to some ions. The absorption of these ions is facilitated by some special proteins on the plasma membrane called Carrier Proteins . First, the carrier proteins are combined with the ions to form the carrier-ion- complex. The carrier-ion-
complex can move across the plasma membrane. When the carrier- ion-complex reaches the inner surface of the membrane, they release the ions into the lumen of the cell. After this, the carrier protein will go back to the outer surface to accept new ions. (3). Donnan Equilibrium The Donnan equilibrium explains the accumulation of some mineral ions inside the cell against the concentration gradient without the expenditure of metabolic energy. According to this theory, inside the root cells, there are some ions called Indiffusable Ions or Fixed Ions, which do not diffuse outside through the plasma membrane. The plasma membrane is permeable to other type of cations and anions. If a cell has an accumulation of cations as fixed ions, such a cell can absorb anions from the soil in order to maintain the electrical potential balance. The absorption of anion in this case causes an equilibrium of both cations and anion inside the cell. This equilibrium is called Donnan equilibrium. By attaining the Donnan equilibrium, the root cells can absorb any minerals in their ionic form.
